Making the Big Trip Across the USA

My internship with the CLM program is beginning to come to an end. I have had great experiences out in Nevada that will help prepare me for the next step in my career. I have met some amazing people who have turned out to be good friends and I plan on keeping in touch with them even after we’re separated by time and distance.

My original interest in the CLM program was sparked by many of the benefits listed on their web site and I can confidently say that I did receive the benefits that I was so enticed by in the beginning of this journey. I have learned about working at a federal agency, my resume has expanded, I was able to explore new habitats and landscapes more intimately then I first even imagined, and I felt proud to be working in the environmental sector soon after graduation. My 7 months with the CLM has been quite the adventure.

So now it’s time to start packing up my things and saying my goodbyes. I’m excited to go back home and be close to friends and family for the holidays but I have a feeling I’ll always feel a pang of nostalgia when I think of my time spent out west.

Adios! Farewell! Ciao! I bid you all adieu!

Intervarietal Promiscuity At El Capitan

Erigonum fasciculatum at El Capitan Open Space preserve

This internship has provided a lot of new experiences and opportunities for learning. Specifically, events during the collection of Eriogonum fasciculatum seeds at El Capitan open space preserve compelled me to review some basic biological concepts regarding speciation.
The whole exercise began when Lauren (my internship colleague) and I went on a seemingly routine mission to collect Eriogonum seeds. By the time we each filled a quarter of a bucket with seeds and associated material, Lauren pointed out extensive variation in the leaf morphology of the plants we were collecting. Some plants had linear light green leaves while others had much wider, blue green, tomentose leaves. To make matters more confusing, some plants had shoots exhibiting both leaf types. This made us think there was hybridization of some sort going on so we decided to do some further research before packaging this collection in order to ensure we had taxonomically uniform specimens.
Upon consulting the natural history museum’s plant atlas, we learned that there are three varieties of Eriogonum fasciculatum in the area we were sampling. These are var. fasciculatum, var. foliolosum, and var. polifolium. According to the Jepson manual, var. polifolium has the wider, oblanceolate, tomentose leaves while var. foliolosum has the linear leaves which aren’t nearly as wooly on their upper surface. While var. faciculatum also occurs in the area, I didn’t find clear examples like I did the other varieties. Consequently, we inferred that the individuals we encountered were vars. foliolosum, polifolium, and mixes thereof.
Although we determined the collection was not useful to us because we needed seeds whose varietal lineage was free of co-mingling, this left me with some fundamental questions. How come these morphologically distinct populations are not classified as different species ? What is the difference between a species and a variety ?
Lauren Riesberg, in her extensive writings addressing speciation and hybridization, cited Mayr’s definition of a species as a “groups of interbreeding natural populations reproductively isolated from other such groups”. However, species that are not geographically isolated have been known to interbreed and produce hybrid offspring. Wikipedia defines a variety as a genetically, and in turn, morphologically distinct subset of a species that is geographically isolated from other populations within that species. However, when the geographic barrier is removed, this subset will interbreed with the rest of the species, resulting in an influx of genes which will erode the variety’s distinct features, thus reintegrating it into the greater species group. With this in mind, I distinguish species and varieties based on the fact that a hybrid of two species will produce offspring with reduced fertility, such as malformed pollen grains, while a hybrid of two subspecific varieties will produce offspring that show no such deformities and will reflect the traits of the original species. This concept reflects the methodology Michael Mesler uses in his assessment of hybridity.
In light of these considerations, I came to the conclusion that El Capitan open space preserve is a very special place in the origins and family history of Eriogonum fasciculatum . According to Flora of North America, Eriogonum fasciculatum var. folilosum came about from an ancient hybridization event between vars. fasciculatum and polifolium. This event likely happened at a place much like El Capitan open space reserve, where these populations intermingle. Eventually, isolated populations of these varieties might inbreed to the point that they become reproductively isolated from the parent varieties, giving rise to new species through divergent evolution. El Capitan offers a living snapshot into the evolutionary history that drives speciation. In conclusion, this experience reminds me that speciation is a dynamic process taking place in our backyards, in present times, while we go about our daily lives.

-Miguel K., Escondido, CA

Desert Trip

Pretty much from day one at the institute I’ve heard people talk about how beautiful Anza-Borrego Desert is. This year’s Seeds of Success collections focused more on the areas surrounding Ramona. While the properties we scouted in the Ramona area were very diverse and interesting, they were all composed of the same types of plant communities. This was the first week we’ve been able to make it out to the desert transition areas leading into Anza-Borrego. When planning to explore a new property, the first step is to find one that has access. For our internship, we are only allowed to make collections on BLM land, which doesn’t always have roads. We look at layers of BLM land on GIS and then choose one that looks reasonable. To do this we mostly use images from Google Earth. The only problem with this is that it can be difficult to determine if something is a dry creek or a road… but we were fairly sure that we could get within walking distance of the site. Another part of scouting that is always fun is navigating back roads. The first dirt road we turned off on was well kept, the second one was less so, and on the third road I felt the tires sink a few inches into soft ground. We managed not to get the truck stuck, which was a very good thing since we didn’t have cell reception. The site itself was a combination of hills and dry wash ravines with an amazing number of different plant species. After exploring for a couple of hours we determined that we could make three collections starting next week, which was really exciting!